Corynexochida

Damesella paronai

Order Corynexochida / Suborder Leiostegiina / Superfamily Dameselloidea / Family Damesellidae / Subfamily Damesellinae

Guzhangian (Miaolingian, Cambrian)Shandong and other parts of North China

Damesella paronai — photo 1 — Photo 1 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 2 — Photo 2 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 3 — Photo 3 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 4 — Photo 4 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 5 — Photo 5 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 6 — Photo 6 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 7 — Photo 7 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 8 — Photo 8 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 9 — Photo 9 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Damesella paronai — photo 10 — Photo 10 · Lectotype (Shandong; photograph courtesy of Restoration Prof. Peng Shanchi) (illustrated by Haixiang) Complete dorsal exoskeleton Complete dorsal exoskeleton Complete dorsal exoskeleton Ventrally compressed specimen (Shandong; Baishui) (Shandong; ilaoniu) (Shandong; Mr. Heiyanquan) (Shandong; Pengcheng Jianke) Specimens showing healed Malformed specimen with injury (left) and malformed reduced spines and fused median pygidial spines (right) pygidial spines (Shandong; (Shandong; Dai Xu) ilaoniu)

Classification

Order Corynexochida / Suborder Leiostegiina / Superfamily Dameselloidea / Family Damesellidae / Subfamily Damesellinae

Genus

Damesella Walcott, 1905

Genus author

Walcott C. D.

Genus etymology

Dames, an early researcher of Chinese trilobites

Species

D. paronai (Airaghi, 1902)

Species author

Airaghi C.

Species etymology

Parona, a surname

Age

Guzhangian (Miaolingian, Cambrian)

Distribution

Shandong and other parts of North China

Synonyms

Cheirurus paronai; Damesella blackwelderi; D. bellgranulata; D. walcotti; D. manchuriensis; D. leishuiensis

Diagnosis

Entire dorsal shield covered with tubercles of varying size. Cranidium trapezoidal; glabella truncate-conical with rounded front; three pairs of glabellar furrows, the anterior two pairs shallower; occipital ring convex, of uniform width, occipital furrow deep; preglabellar field absent; palpebral lobes short to medium; eye ridges distinct; librigenae with wide genal field and stout, medium-length genal spines. Thorax of 12 segments. Pygidium subtriangular to semicircular; axis long with rounded end; 6 pairs of marginal spines, pairs 2 and 6 shorter.

Remarks

This species is the type species of the genus Damesella. Kobayashi (1941) designated Airaghi's (1902) enrolled specimen as the type, an action invalidated by Yuan Jinliang et al. (2012).